2026 in paleomalacology

This list records new taxa of fossil molluscs that were announced or described in 2026. Other peer-reviewed publications on discoveries related to paleomalacology which occurred in that year are also detailed here.

Ammonites

Name Novelty Status Authors Age Type locality Country Notes Images

Eckhardites atmensis[1]

Sp. nov

Valid

Mitta in Mitta, Zenina & Meleshin

Middle Jurassic (Bathonian)

Russia

A member of the family Cardioceratidae. Published online in 2026, but the issue date is listed as December 2025.

Leonardia[2]

Gen. et sp. nov

Parent & Garrido

Jurassic-Cretaceous transition

Vaca Muerta

Argentina

Genus includes new species L. almanzaensis. The generic name is shared with Leonardia Canefri (1890) and Leonardia Mearns (1905).

Ammonite research

  • Neige & van Tiel (2026) study the evolutionary history of the family Dactylioceratidae, and interpret the pattern of changes of their taxonomic and morphological diversity as consistent with background extinction, possibly linked to species specialization making them vulnerable to such extinction.[3]
  • Aguirre-Urreta et al. (2026) report evidence of exceptional preservation of the periostracum in specimens of Bochianites neocomiensis and Lissonia riveroi from Vaca Muerta (Argentina), interpreted as indicative of highly conservative composition of the periostracum throughout the evolutionary history of conchiferan molluscs.[4]
  • Frau (2026) revises the affinities of the genera Vergunniceras and Paracheloniceras, and names a new subfamily Paracheloniceratinae within the family Douvilleiceratidae.[5]
  • A study on the biostratigraphy of the Barremian-Aptian ammonite fossils from the southern edge of the Vercors Urgonian platform (France) is published by Pictet, Ferry & Pietra (2026).[6]
  • Hefny et al. (2026) revise the composition of Cenomanian and Turonian ammonite assemblages from the strata of the Galala and Umm Omeiyid formations at Wadi Qena (Egypt).[7]
  • Bensekhria et al. (2026) study the biostratigraphy of the Albian–Turonian ammonite fossils from the Aurès Basin (Algeria) and compare the composition of the studied ammonite assemblages with those from Tunisia, Western Europe and the Western Interior Seaway, reporting evidence of differences that were likely driven by climatic gradients, marine barriers and variable migration pathways.[8]

Other cephalopods

Name Novelty Status Authors Age Type locality Country Notes Images

Discoceras alienum[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Katian)

Germany

Discoceras angustum[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Sandbian or Katian)

Estonia

Discoceras aseriense[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Darriwilian)

Estonia

Discoceras brandenburgense[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Katian)

Germany

Discoceras circulare[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Sandbian)

Estonia
Germany

Discoceras polonicum[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Katian)

Poland

Discoceras sovaki[9]

Sp. nov

Valid

Aubrechtová, Korn & Kröger

Ordovician (Katian)

Germany

Uluciala[10]

Gen. et sp. nov

Valid

Sugiura et al.

Late Cretaceous (Campanian-Maastrichtian)

United States
( South Dakota)

A member of Decabrachia of uncertain affinities, with a morphology intermediate between those of members of Sepiida and Sepiolida. The type species is U. rotundata.

Other cephalopod research

  • Evidence of impact of distribution of mineral deposits in chambered shells on buoyancy and maneuverability of orthocone cephalopods is presented by Peterman, Landman & Ciampaglio (2026).[11]
  • Patarroyo et al. (2026) describe new fossil material of Aturia peruviana from the Paleogene strata of the San Jacinto Formation (Colombia), and interpret their morphology as supporting the classification of Aturia colombiana as a junior synonym of A. peruviana.[12]
  • Evidence from the study of extant nautiloid cephalopods and from the study of oxygen isotope shell thermometry of nautiloid fossils, indicating that modern nautiloids live deeper and grow in colder water than any of their extinct relatives other than members of the genus Aturia, is presented by Ward et al. (2026).[13]

Bivalves

Name Novelty Status Authors Age Type locality Location Notes Images

Bicornucopina acuminata[14]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Barremian-Aptian)

France

A member of the family Monopleuridae.

Cyathodonta gilardeti[15]

Nom. nov

Valid

Pacaud in Pacaud et al.

Eocene

France

A species of Cyathodonta; a replacement name for Thracia rugosa Bellardi (1852).

Cyathodonta tarbelliana[15]

Sp. nov

Valid

Pacaud et al.

Eocene

Brassempouy Limestone

France

A species of Cyathodonta.

Homopleura parva[14]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Monopleuridae.

Horiopleura arabica[16]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

Qishn Formation

Oman

A member of the family Polyconitidae.

Horiopleura helviorum[16]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Polyconitidae.

Hypelasma fritzlangi[17]

Sp. nov

Valid

Schneider & Werner

Late Jurassic (Kimmeridgian)

Frankenalb Formation

Germany

A rudist bivalve belonging to the family Requieniidae.

Mazaevella[18]

Gen. et comb. nov

Valid

Shilekhin, Biakov & Vdovichenko

Carboniferous-Permian (Gzhelian-Sakmarian)

Russia

A member of Pectinida belonging to the family Annuliconchidae. The type species is "Annuliconcha" placunensis Nelzina (1958). Published online in 2026, but the issue date is listed as December 2025.

Myostyla[14]

Gen. et sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Monopleuridae. The type species is M. ardescensis.

Praecaprina paquieri[14]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Caprinidae.

Praecaprina tubifera[14]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Caprinidae.

Praecaprina vacua[14]

Sp. nov

Valid

Masse et al.

Early Cretaceous (Aptian)

France

A member of the family Caprinidae.

Saalidiceras[17]

Gen. et sp. nov

Valid

Schneider & Werner

Late Jurassic (Kimmeridgian)

Frankenalb Formation

Germany

A rudist bivalve belonging to the family Epidiceratidae. The type species is S. syllai.

Thracia sixi[15]

Sp. nov

Valid

Pacaud et al.

Eocene

Brassempouy Limestone

France

A species of Thracia.

Valclusella[14]

Gen. et 2 sp. nov

Valid

Masse et al.

Early Cretaceous (Barremian-Aptian)

France

A member of the family Monopleuridae. The type species is V. valclusensis; genus also includes V. biconvexa.

Bivalve research

  • Evidence of utility of the study of rudist shells from the Maastrichtian Cárdenas Formation (Mexico) in reconstruction of environmental conditions in the western Tethys Ocean during the Cretaceous is presented by Juárez-Aguilar et al. (2026).[19]
  • Pérez, Mosquera & Cuitiño (2026) report the discovery of fossil material of Limopsis insolita from the Miocene Gaiman Formation (Argentina), extending its known geographic distribution northwards and representing the first unambiguous record of a member of the genus Limopsis in deeper-water settings in the region.[20]
  • Knight (2026) studies the composition of two assemblages of Miocene (Tortonian) bivalves from S'Algar (Menorca, Spain), interpreted as originating from a middle ramp environment that changed through time as a result of changes in the frequency and directionality of storms and currents.[21]

Gastropods

Name Novelty Status Authors Age Type locality Location Notes Images

Acrostemma anatolica[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Turkey

A member of the family Cylichnidae.

Acteon antipinguis[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Acteon.

Acteon gutta[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Acteon.

Acteon romanicus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Acteon.

Acteon trifasciatus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Acteon.

Acteon viciani[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Hungary

A species of Acteon.

Acteon vindobonensis[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Acteon.

Acteonella murcielaguensis[23]

Sp. nov

Valid

Aguilar et al.

Late Cretaceous (Maastrichtian)

El Viejo Formation

Costa Rica

Angulariopsis[24]

Gen. et comb. nov

Valid

Monari et al.

Late Triassic (Rhaetian) and Early Jurassic (Hettangian)

Austria
France
Luxembourg

A member of the family Purpurinidae. The type species is "Angularia" nivernensis Dareste de la Chavanne (1912); genus also includes "Angularia" corallina Nützel et al. (2022).

Bittium biskupici[25]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

Austria

A species of Bittium.

Bittium castori[25]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

Romania

A species of Bittium.

Bittium pallgergelyi[25]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Baden Formation

Austria

A species of Bittium.

Bittium parvapyramis[25]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Ukraine

A species of Bittium.

Bittium polluxi[25]

Sp. nov

Valid

Harzhauser & Landau

Miocene

Dej Formation

Romania

A species of Bittium.

Bourguetia bipartita[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Pseudomelaniidae.

Brouchilda[24]

Gen. et 2 sp. et comb. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

France
Luxembourg

A member of the family Mathildidae. The type species is B. laevigata; genus also includes new species B. mulleri, as well as "Promathildia" bicarinata Dareste de la Chavanne (1912).

Bulla? elalbanii[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Ukraine

Possibly a species of Bulla.

Campanile metaisi[26]

Sp. nov

Valid

Merle & Pacaud in Merle et al.

Eocene

Lakhra Formation

Pakistan

A species of Campanile.

Campanistylus[26]

Gen. et sp. nov

Valid

Merle & Pacaud in Merle et al.

Eocene

Lakhra Formation

Pakistan

A member of the family Campanilidae. The type species is C. lakhraensis.

Chelidonura piseraii[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Poland

A species of Chelidonura.

Conotomaria stoliczkai[27]

Sp. nov

Ghosh et al.

Late Cretaceous

India

A member of the family Pleurotomariidae.

Cylichna caudex[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Cylichna.

Cylichna dolium[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Czech Republic

A species of Cylichna.

Cylichna eschnerae[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Czech Republic

A species of Cylichna.

Cylichna miosimplex[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Cylichna.

Cylichna steiningeri[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Cylichna.

Cylichna straussi[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Cylichna.

Dakshinatomaria[27]

Gen. et sp. nov

Ghosh et al.

Late Cretaceous

India

A member of the family Pleurotomariidae. Genus includes new species D. rajendrai.

Globularia delsatei[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

Gruendelifusus[24]

Gen. et sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A possible member of the superfamily Mathildoidea. The type species is G. melusinae.

Jurazyga[24]

Gen. et comb. nov

Valid

Monari et al.

Early Jurassic (Hettangian and Pliensbachian)

France
Italy
Luxembourg

A member of the family Zygopleuridae. The type species is "Melania" theodori Terquem (1855); genus also includes "Zygopleura" subnodosa (d'Orbigny 1850), "Zygopleura" vinosimonensis Fischer & Weber (1997), "Chemnitzia" polyplecta Gemmellaro (1878), "Chemnitzia" moorei Gemmellaro (1878), "Chemnitzia" appenninica Gemmellaro (1878) and "Chemnitzia" veturia Gemmellaro (1878).

Kaimella[24]

Gen. et sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A possible member of the family Cimidae. The type species is K. tenuilineata.

Microschiza pauciornata[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Purpuroideidae.

Miracypraea[28]

Gen. et sp. nov

Valid

Pacaud, Gómez-García & Celzard

Eocene

Spain

A member of the superfamily Cypraeoidea belonging to the family Eocypraeidae. The type species is M. alluensis.

Neubaueriella[22]

Gen. et comb. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A member of the family Haminoeidae; a new genus for "Atys (Alicula)" lapugyensis Berger (1949).

Oonia feidtorum[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Ampullinidae.

Oonia haasi[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Ampullinidae.

Pseudavena bella[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Czech Republic

A member of Heterobranchia of uncertain affinities.

Pyrunculus microromanicus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A member of the family Retusidae.

Pyrunculus szobiensis[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Hungary

A member of the family Retusidae.

Pyrunculus zuschini[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A member of the family Retusidae.

Retusa mystica[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Retusa.

Rictaxis austriacus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Rictaxis.

Rictaxis siedli[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Rictaxis.

Rictaxis ursulae[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Rictaxis.

Ringicula daisyae[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Ringicula.

Roxania bomba[22]

Nom. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Roxania; a replacement name for Sabatia callifera helvetica Berger (1949).

Roxania oviformis[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Roxania.

Roxania pila[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Roxania.

Sabatia quasimodoi[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Czech Republic

A species of Sabatia.

Scaphander alatissimus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Scaphander.

Scaphander catena[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Romania

A species of Scaphander.

Scaphander perforatus[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Austria

A species of Scaphander.

Scaphander schollnbergeri[22]

Sp. nov

Valid

Harzhauser, Landau & Malaquias

Miocene

Czech Republic

A species of Scaphander.

Tangarilda darestei[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Mathildidae.

Turritelloidea? bockfielsensis[24]

Sp. nov

Valid

Monari et al.

Early Jurassic (Hettangian)

Luxembourg Sandstone

Luxembourg

A member of the family Gordenellidae.

Gastropod research

  • Evidence of a southward expansion of nerineoid gastropods from the Boreal Realm into the Tethyan Realm during the Jurassic period is presented by Leshno Afriat, Rabinovich & Edelman-Furstenberg (2026).[29]
  • Li, Xiao & Yu (2026) report the discovery of new fossil material of Coptocheilus electrothauma (originally named Schistoloma electrothauma) from the Cretaceous amber from Myanmar, and revise the diagnostic traits of this species.[30]

General research

  • Bellosi et al. (2026) revise the age and distribution of the molluscan assemblages from Patagonia living at the time of the Chattian-Langhian marine incursions into southern South America, and provide calibrated dating of the youngest fauna dominated by tropical species.[31]
  • DeVries (2026) studies the composition of late Pliocene and Pleistocene marine molluscan assemblages from northwestern Peru, and reports evidence of a shift from a cool-water to warm-water fauna during the Pleistocene that might have been linked to uplift of the coastal plain and/or altered equatorial circulation.[32]
  • Betz et al. (2026) evaluate functional traits of Pliocene-Holocene bivalves and gastropods from the West Atlantic, finding no evidence of a significant association of the studied traits with increased extinction risk of the studied molluscs.[33]

References

  1. ^ Mitta, V. V.; Zenina, Ju. V.; Meleshin, I. A. (2026). "First Finds of Eckhardites (Ammonoidea: Cardioceratidae) in the Upper Bathonian of the Russian Platform". Paleontological Journal. 59 (6): 620–628. doi:10.1134/S0031030125600982.
  2. ^ Parent, H.; Garrido, A. C. (2026). "The Upper Tithonian–Lower Berriasian ammonite succession of Cajón de Almanza (Vaca Muerta Formation, Neuquén Basin, Argentina)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 316 (3): 347–365. doi:10.1127/njgpa/1299.
  3. ^ Neige, P.; van Tiel, T. (2026). "Background extinction due to species specialization? Insights from a high-resolution Jurassic ammonoid case study (Dactylioceratidae)". Paleobiology: 1–17. doi:10.1017/pab.2025.10086.
  4. ^ Aguirre-Urreta, B.; Marin, L. S.; Checa, A. G.; Grenier, C.; Tunik, M.; Lescano, M.; Castro, M. A.; Lazo, D. G.; Vennari, V. V.; Rogel, M. N. (2026). "Organic periostracum preserved in Cretaceous ammonoids from the Andean Neuquén Basin". Communications Biology. doi:10.1038/s42003-026-09635-6.
  5. ^ Frau, C. (2026). "Taxonomy and significance of the Aptian ammonite complex Vergunniceras Thomel, 1980 and Paracheloniceras Collignon, 1962, and a new subfamily Paracheloniceratinae". Palaeoworld 201079. doi:10.1016/j.palwor.2026.201079.
  6. ^ Pictet, A.; Ferry, S.; Pietra, L. (2026). "Ammonite biostratigraphy on the platform–slope transition between the Vercors Urgonian platform and the Vocontian Trough (S–E France)". Swiss Journal of Palaeontology. 145: 17–76. doi:10.3897/sjp.145.175636.
  7. ^ Hefny, A.; El Qot, G.; El Araby, A.; Aly, M. F. (2026). "Cenomanian–Turonian ammonites from Wadi Qena, Eastern Desert, Egypt: systematic paleontology and biostratigraphy". Journal of African Earth Sciences 105996. doi:10.1016/j.jafrearsci.2026.105996.
  8. ^ Bensekhria, A.; Abdeldjalil, M.; Bouhata, R.; Khelali, M. (2026). "Toward a refined framework for upper Albian-lower Turonian ammonite biostratigraphy: case study from the Aurès Basin (NE Algeria), and regional correlation with Western Europe, central Tunisia, and the Western Interior Seaway". Journal of African Earth Sciences. 237 106046. doi:10.1016/j.jafrearsci.2026.106046.
  9. ^ a b c d e f g Aubrechtová, M.; Korn, D.; Kröger, B. (2026). "The genus Discoceras (Tarphyceratida, Cephalopoda; Ordovician) from Estonia and glacial erratics in north-central Europe". European Journal of Taxonomy. 1036: 1–54. doi:10.5852/ejt.2026.1036.3173.
  10. ^ Sugiura, K.; Ikegami, S.; Takeda, Y.; Mutterlose, J.; Derin, M. O.; Kubota, A.; Nishida, H.; Tainaka, K.; Harada, T.; Landman, N. H.; Iba, Y. (2026). "The oldest sepioid cephalopod from the Cretaceous discovered by Digital fossil-mining with zero-shot learning AI". Communications Biology. doi:10.1038/s42003-026-09519-9. PMID 41545472.
  11. ^ Peterman, D. J.; Landman, N.; Ciampaglio, C. (2026). "Exploring the influence of cameral deposits on the stability, orientation, and maneuverability of orthocone cephalopods". Paleobiology: 1–21. doi:10.1017/pab.2025.10085.
  12. ^ Patarroyo, P.; Casallas, W.; Benavides-Cabra, C. D.; Herrera, J. M. (2026). "Records of Aturia peruviana (Nautilida, Cephalopoda) from the San Jacinto Formation (upper Eocene to lower Oligocene), Ovejas (Sucre, Colombia)". Publicación Electrónica de la Asociación Paleontológica Argentina. 26 (1): 37–46. doi:10.5710/PEAPA.20.10.2025.549.
  13. ^ Ward, P. D.; Barord, G.; Carlson, B.; Dooley, F.; Dunstan, A.; Gowep, N.; Hamilton, R.; Heuring, W.; Kinch, J.; Klug, C.; Landman, N.; Schauer, A.; Steig, E. J.; Veloso, J. (2026). "Comparative habits and habitat in extant and extinct nautiloid cephalopods from acoustic telemetry and stable oxygen isotope analyses". Scientific Reports. doi:10.1038/s41598-026-36623-x.
  14. ^ a b c d e f g Masse, M.; Rineau, V.; Fenerci-Masse, M.; Frau, C. (2026). "Upper Barremian-lower Aptian rudist faunas of Urgonian-type platform formations from Ardèche (southeastern France)". Carnets Geol. 26 (3): 51–87. doi:10.2110/carnets.2026.2603.
  15. ^ a b c Pacaud, J.-M.; Audouit, P.; Coppini, S.; Buisson, S.; Meunier, F. (2026). "New Cyathodonta and Thracia species (Bivalvia, Thraciidae) from the Bartonian (Middle Eocene) of Horsarrieu [Pédelail] (Landes, France)". Acta Malacologica Inquisitionis. 2 (2): 114–121.
  16. ^ a b Masse, M.; Fenerci-Masse, M.; Platel, J.-P.; Frau, C.; Léonide, P. (2026). "New species of the genus Horiopleura (Bivalvia, Hippuritida, Polyconitidae) from the Barremian-lower Aptian deposits of the Mediterranean region: evolutionary implications". Acta Palaeontologica Romaniae. 21 (2): 41–52. doi:10.35463/j.apr.2025.02.06.
  17. ^ a b Schneider, S.; Werner, W. (2026). "The late Kimmeridgian bivalve fauna of Saal an der Donau (Bavaria, southern Germany): the Hippuritida". Zitteliana. 100: 1–30. doi:10.3897/zitteliana.100.169922.
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